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Butterfly taxonomy (classification) The skippers The swallowtails The whites The hairstreaks, coppers and blues. Includes the Duke of Burgundy. The nymphalids, fritillaries and browns. Includes the Monarch.
Wall - imago - Arnside Knott - 07-Aug-06 (0584)
Wingspan
45 - 53mm
Photo © Pete Eeles
Wall

Lasiommata megera
Pronunciation LASS-ee-oh-may-tuh
muh-JEE-ruh
B&F No.: 1615
Superfamily: Papilionoidea
Family: Nymphalidae
Subfamily: Satyrinae
Genus: Lasiommata
Subgenus:  
Species: megera (Linnaeus, 1767)
Subspecies:megera (Linnaeus, 1767)
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Introduction  

The Wall gets its name from the characteristic behaviour of resting with wings two-thirds open on any bare surface, including bare ground and, of course, walls! Many people will have come across this butterfly on footpaths, especially in coastal areas, where the butterfly flies up when disturbed, before setting again a few metres ahead.

The basking behaviour of this butterfly allows it to benefit from the full warmth of the sun whose rays shine directly on the butterfly, but also get reflected back onto the butterfly from whichever surface it is resting on. This habit allows the butterfly to raise its body temperature sufficiently high for it to fly. In particularly hot weather, however, such basking is avoided and the butterfly may even retreat to a suitably-shaded spot to avoid overheating.

This species was once found throughout England, Wales, Ireland and parts of Scotland. Today, however, is a very different picture, with this species suffering severe declines over the last several decades. It is now confined to primarily-coastal regions and has been lost from many sites in central, eastern and south-east England. In Scotland it is confined to coastal areas in the south-west of the country. It is also found on the Isle of Man and Channel Islands. This butterfly is found in relatively small colonies that are self-contained although some individuals will wander, allowing the species to quickly colonise suitable nearby sites.

Subspecies: Lasiommata megera megera

The nominate subspecies was first defined in Linnaeus (1767) as shown here (type locality: Austria and Denmark). The population in the British Isles is represented by this subspecies.


Wall Brown, 12/08/2012, Steyning
Male
Photo © Pauline
Wall Brown, male. Seaford, East Sussex. 24/5/2012.
Male Underside
Photo © badgerbob
Wall female- Bonchurch Down Isle of Wight-15 May 2012
Female
Photo © marmari
Wall Female - Ouse Washes, Cambs - 30.07.2010
Female Underside
Photo © PhiliB

Phenology  

The first generation of adults emerge in early May, peaking at the end of May and early June, or a little later in the north of England and Scotland. They give rise to a second brood that emerges at the end of July, or mid-August further north. There are 2 generations each year and, on occasion, a small 3rd generation may appear in October.

Subspecies: Lasiommata megera megera


The chart(s) above have been correlated with the phenology plot below, taken from the UK Butterfly Monitoring Scheme. The blue line gives average counts over the full data set from 1976 to date, and the red line gives the average for the last year.


Habitat  

This species is now found primarily in coastal areas, especially unimproved grassland, wasteland, cliff edges and hedgerows.

Larval Foodplants  

The primary larval foodplants are Bents (various) (Agrostis spp.), Cock's-foot (Dactylis glomerata), False Brome (Brachypodium sylvaticum), Tor-grass (Brachypodium pinnatum), Wavy Hair-grass (Deschampsia flexuosa) and Yorkshire-fog (Holcus lanatus).

Nectar Sources  

Adults feed primarily on Daisy (Bellis perennis), Fleabane (Pulicaria dysenterica), Hawkweeds (Hieracium/Hypochoeris), Knapweeds (Centaurea spp.), Marjoram (Origanum vulgare), Ragged Robin (Lychnis flos-cuculi), Ragwort (Senecio jacobaea), Thistles (Cirsium spp. and Carduus spp.), Water Mint (Mentha aquatica) and Yarrow (Achillea millefolium).

Imago  

The male of this species is territorial and will inhabit a particular area, such as a path, hedgerow or roadside verge, waiting for a passing female. Males will typically perch in a favoured position but will, in sunny and warm conditions, adopt a strategy of patrolling in order to find a mate. All passing insects are investigated and rival males will fly high into the air before coming back to the ground a few seconds later.

The female is much more sedentary and the less-conspicuous of the two sexes. After a brief courtship a pair will mate before disappearing into surrounding vegetation. Both sexes are avid nectar feeders and will feed from any available flower.

Subspecies: Lasiommata megera megera


Wall pair - Durlston Country Park, Dorset 27-May-05
Photo © Vince Massimo
27-May-2005
Wall Male - Ballard Down, Dorset 22-May-05
Photo © Vince Massimo
22-May-2005
Wall - imago - Arnside Knott - 07-Aug-06 (0584)
Photo © Pete Eeles
07-Aug-2006
Wall - imago - Arnside Knott - 07-Aug-06 (0585)
Photo © Pete Eeles
07-Aug-2006
Wall - imago - Arnside Knott - 08-Aug-07 (1140)
Photo © Pete Eeles
08-Aug-2007
Wall - imago - Thatcham - 16-May-05 [REARED]
Photo © Pete Eeles
17-May-2005
Wall, Steyning, 10 August 2008
Photo © Sussex Kipper
10-Aug-2008
Wall Female - Ouse Washes, Cambs - 30.07.2010
Photo © PhiliB
30-Jul-2010
Wall Female - Crawley, Sussex 9-Oct-08
Photo © Vince Massimo
09-Oct-2008
Wall Female - Crawley, Sussex 9-Oct-08
Photo © Vince Massimo
09-Oct-2008
Wall Brown, male. Seaford, East Sussex. 23/5/2012.
Photo © badgerbob
23-May-2012
Wall, Female. Offham, Lewes. 12/8/2011.
Photo © badgerbob
12-Aug-2011
Mating Wall. 3rd brood. Seaford. 23/9/2011.
Photo © badgerbob
23-Sep-2011
Wall (mating couple) - Alpes-Maritimes - 1 June 2012
Photo © CFB
01-Jun-2012
Wall, Female, Dunglass, East Lothian, 24/08/2010
Scotland
Photo © NickMorgan
Wall, Male, Dunglass, East Lothian, 24/08/2010
Scotland
Photo © NickMorgan
Wall Brown, male. Seaford, East Sussex. 24/5/2012.
Photo © badgerbob
24-May-2012
Wall - imago - Thatcham - 27-May-12 (1) [REARED]
Photo © Pete Eeles
Wall - imago - Thatcham - 27-May-12 (2) [REARED]
Photo © Pete Eeles
Wall, female - Mill Hill, West Sussex 12-August-12
Photo © Colin Knight
12-Aug-2012
Wall Brown, 12/08/2012, Steyning
Photo © Pauline
12-Aug-2012
Wall Brown, 12/08/2012, Steyning
Photo © Pauline
12-Aug-2012
Warren Hills 28-Aug-2010
Photo © steveb
Wall female- Bonchurch Down Isle of Wight-15 May 2012
Photo © marmari
Wall male - Chantry Hill, Sussex 5-Aug-2010
Photo © Sussex Kipper
05-Aug-2010
Wall male - High & Over, Sussex 29-Sept-2011
Photo © Sussex Kipper
29-Sep-2011
Wall female - Steyning, Sussex 6-Aug-2010
Photo © Sussex Kipper
06-Aug-2010

Aberrations  

Description to be completed.

Unclassified Aberrations


Wall - aberration - Cley next the Sea - 30-May-00 [Alan Roe]
Photo © Alan Roe
wall aberation 31-May-2010
Photo © ColinC
31-May-2010
Wall male (aberration) Castle Hill NNR, East Sussex 30-April-2012
Photo © Average Wingspan
31-May-2012

Ovum  

The spherical eggs are laid singly, or occasionally in twos and threes, in various positions, including the leaves of the foodplant, exposed roots and nearby vegetation. Eggs are pale green when first laid, becoming more translucent as the larva develops within. Sites for egg-laying are typically sheltered and warm compared to their surroundings, and include grass clumps, rabbit scrapes and hoof prints from cattle. This stage lasts around 10 days.


Wall - ovum - Unknown location - 2004 (2) [REARED] [Reg Fry]
Photo © Reg Fry
Wall - ovum - Unknown location - 2004 [REARED] [Reg Fry]
Photo © Reg Fry

Larva  

After emerging, the young larva typically eats its eggshell before feeding on the leaves of the foodplant. Larvae become more mobile as they mature and will move from plant to plant as needed. Larvae typically feed at night, but occasionally feed during the day. This stage lasts around 4 weeks for those larvae that do not overwinter, and there are 3 moults in total.


Location of wall larva, High and Over, Sussex. 31/1/2013.
Photo © badgerbob
31-Jan-2013
Wall larva. High and Over, Sussex. 31/01/2013.
Photo © badgerbob
31-Jan-2013
Wall - larva - Thatcham - 24-Apr-05 (3) [REARED]
Photo © Pete Eeles
24-Apr-2005
Wall - larva - Thatcham - 24-Apr-05 (4) [REARED]
Photo © Pete Eeles
24-Apr-2005
Wall - larva - Thatcham - 24-Apr-05 [REARED]
Photo © Pete Eeles
15-Apr-2005
Wall - larva - Unknown location - 2004 (2) [REARED] [Reg Fry]
Photo © Reg Fry
Wall - larva - Unknown location - 2004 (3) [REARED] [Reg Fry]
Photo © Reg Fry
Wall - larva - Unknown location - 2004 [REARED] [Reg Fry]
Photo © Reg Fry
Wall Brown larvae. High and Over, Sussex. 30/01/2013.
Approx 1.5cm length.
Photo © badgerbob
30-Jan-2013
Wall larva. High and Over, Sussex. 31/01/2013.
Photo © badgerbob
31-Jan-2013
Wall Brown Larva - High and Over, Seaford, Sussex 20-March-2012 (DSC7864)
Photo © badgerbob
20-Mar-2012
Wall larva. High and Over, Seaford. 20/3/2012
Photo © badgerbob
20-Mar-2012
Wall larva, High and Over, Seaford. 28/3/2012.
Photo © badgerbob
28-Mar-2012
Wall Brown larva following moult. 6/3/2013. High and Over. Seaford.
Photo © badgerbob
06-Mar-2013
Moulting Wall Brown larva. 5/3/2013 High and Over. Seaford.
Photo © badgerbob
Moulting Wall Brown larva. 5/3/2013 High and Over. Seaford.
Photo © badgerbob
05-Mar-2013
Wall Brown larva. 18/3/2013. High and Over, Seaford.
Photo © badgerbob
18-Mar-2013
Wall Brown larva. 18/3/2013. High and Over. Seaford.
Photo © badgerbob
18-Mar-2013
Wall Brown larva. High and Over, Seaford. 27/3/2013.
Photo © badgerbob
27-Mar-2013
Wall Brown larva. High and Over. 6/4/2013.
Photo © badgerbob
06-Apr-2013
Wall larva 4 days before pupation. 19/4/2013. High and Over.
Photo © badgerbob
19-Apr-2013
Wall larva 4 days before pupation. 19/4/2013. High and Over. Seaford.
Photo © badgerbob
19-Apr-2013

Pupa  

The green pupa is formed head down, attached by the cremaster to the foodplant or nearby vegetation and is extremely well camouflaged. This stage lasts around 2 weeks.


Wall - pupa - Thatcham - 24-Apr-05 (2) [REARED]
Photo © Pete Eeles
24-Apr-2005
Wall - pupa - Thatcham - 24-Apr-05 [REARED]
Photo © Pete Eeles
24-Apr-2005
Wall - pupa - Unknown location - 2004 (2) [REARED] [Reg Fry]
Photo © Reg Fry
Wall - pupa - Unknown location - 2004 [REARED] [Reg Fry]
Photo © Reg Fry
Wall pupa. High and Over, Sussex 21/4/2012.
Photo © badgerbob
21-Apr-2012
Wall pupa. High and Over, Sussex 20/4/2012.
Photo © badgerbob
20-Apr-2012
Wall Brown pupa. High and Over. Sussex. 19/5/2012.
Photo © badgerbob
19-May-2012

Similar Species  

No similar species found.

Videos  

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Sites  

Click here to see the distribution of this species overlaid with specific site information. Alternatively, select one of the sites listed below.

Sites
Arnside Knott, Badbury Rings, Ballard Down, Ballydyan, Ballyscanlan, Barnack Hills and Holes NNR, Bindon Hill, Bingham Linear Park, Bishop Middleham Quarry, Boultham Mere, Brackett's Coppice, Bradfield Woods, Brean Down, Broadcroft Quarry Reserve, Broaks Wood, Bryncelyn Hall, Candlesby Hill Quarry, Castle Eden Dene, Castlehill Point, Clubmen's Down, Cop Lane, Corfe Castle, Crickley Hill, Crook Peak, Dolebury Warren, Draycott Sleights, Drigg Dunes, Durlston Country Park, Eakring Meadows Nature Reserve, East of Cove Farm, East Ord, Fingringhoe Wick, Fleetwood Marsh, Fontmell Down, Hethfelton Wood, Hexham, Higher Hyde, Hinkley Point Nature Reserve, Hod Hill, Holton Lee, Howth Head, Jerry's Hole, Kenfig Pool, Kingcombe Redholm, Kingcombe Stones, Long Knoll, Lorton Meadows, Lough Bunny, Lough George, Lydlinch Common, Malling Down, Meanwood Park, Mill Hill, Mill Hill Quarry, Monk's Wood, Mynydd Marian, Old Castle Down, Perryfields Quarry, Piddles Wood, Portland Tout Quarry, Powerstock Common, Ringstead Bay, Sand Point, Shipley Station Meadow, Steyning Downs, Stoke Camp, Stubhampton Bottom, Tadnoll, Titley Pool, Tophill Low, Torr Works, Tyne Riverside Country Park, Ubley Warren, Wall Common, Walters Copse, Warton Crag, West Williamston Salt Marshes, Whitbarrow Scar, White Sheet Hill, Windmill Hill and Cleeve Prior, Wiveton Down, Wolla Bank Pit, Wood of Cree, Woolacombe Down

Conservation Status  

There has been a severe and worrying decline of inland populations, with most remaining populations now being found in coastal areas. This species is therefore a priority for conservation efforts.

UK BAP StatusDistribution TrendPopulation Trend
Priority Species
Click here to access the Biodiversity Action Plan (BAP) for this species.
DecreaseDecrease

From The State of Butterflies in Britain and Ireland and the UK Biodiversity Action Plan (UK BAP) (2007 review).


Links  

The following links provide additional information on this butterfly.

References  

The species description provided here has been derived from the author's own observations and the information contained in the following works:

  • Asher (2001). Asher, J., Warren, M., Fox, R., Harding, P., Jeffcoate, G. and Jeffcoate, S.: The Millennium Atlas of Butterflies in Britain and Ireland. 2001.
  • Fox (2006). Fox, R., Asher, J., Brereton, T., Roy, D. and Warren, M.: The State of Butterflies in Britain and Ireland. 2006.
  • Frohawk (1914). Frohawk, F.W.: A Natural History of British Butterflies. 1914.
  • Howarth (1973). Howarth, T.G.: South's British Butterflies. 1973.
  • Nash (2012). Nash, D., Boyd, T. and Hardiman, D.: Ireland's Butterflies: A Review. 2012.
  • Riley (2007). Riley, A.M.: British and Irish Butterflies: The Complete Identification, Field and Site Guide to the Species, Subspecies and Forms. 2007.
  • South (1906). South, R.: The Butterflies Of The British Isles. 1906.
  • Thomas & Lewington (2010). Thomas, J. and Lewington, R.: The Butterflies of Britain and Ireland. Edn.2. 2010.

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