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Butterfly taxonomy (classification) The skippers The swallowtails The whites The hairstreaks, coppers and blues. Includes the Duke of Burgundy. The nymphalids, fritillaries and browns. Includes the Monarch.
Green-veined White male (Second Brood) - Chaldon, Surrey 30-July-2012
Wingspan
40 - 52mm
Photo © Vince Massimo
Green-veined White

Pieris napi
Pronunciation PEE-err-iss
NAY-py
B&F No.: 1551
Superfamily: Papilionoidea
Family: Pieridae
Subfamily: Pierinae
Genus: Pieris
Subgenus:  
Species: napi (Linnaeus, 1758)
Subspecies:sabellicae (Stephens, 1828)
 thomsoni (Warren, 1968)
 britannica (Müller & Kautz, 1939)
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Introduction  

This is a common butterfly of damp grassland and woodland rides and is often mistaken for its cousin, the Small White. It can be found from spring through to autumn in parks and gardens, as well as less-urban areas such as meadows and woodland rides. The so-called green veins on the underside of the adults are, in fact, an illusion created by a subtle combination of yellow and black scales. This is one of the most widespread species found in the British Isles and can be found almost everywhere although it is absent from Shetland and areas of the Scottish Highlands.

Subspecies: Pieris napi napi

The nominate subspecies was first defined in Linnaeus (1758) as shown here (type locality: Sweden). It is not found in the British Isles.

Subspecies: Pieris napi sabellicae

This subspecies was first defined in Stephens (1828) as shown here and as shown in this plate (type locality: England). This subspecies is found in England and Wales. However, in the north of England and southern Scotland, there is no clear dividing line between the distribution of this subspecies and that of the subspecies thomsoni. Bowden (1975) considers the name sabellicae to be redundant, suggesting septentrionalis Verity (1916, type-locality Westcliff-on-sea) instead.

Spring brood males have very feint markings and often lack the single spot on each forewing, whereas summer brood males have more pronounced markings. Spring brood females, on the other hand, are generally more heavily marked than their offspring, especially on the forewings. In both sexes, summer brood adults are generally larger than those of the spring brood.

Pieris napi sabellicae

Allied to Po. Napi, but dissimilar in form, the wings being shorter and more rounded; the anterior being nearly of the form of those of Po. Cardamines,— it has the upper surface of all the wings of a yellowish-white, with broad dusky irrorated nervures; broadest towards the hinder margin : the male has the base of the anterior wings and a single irregular spot in the fourth marginal cell dusky, and the female the base and tips of the same wings, a spot in the fourth and sixth marginal cells, and the inner edge of the wings of the same colour : both sexes have a similarly coloured spot on the upper margin of the posterior wings above. Beneath, all the wings are adorned with very broad dusky nervures, resembling those in var. [zeta: Dilated nervures of the posterior wings beneath dusky in both sexes] of Po. Napi, but varying in different specimens; and the dilated nervure on the upper edge of the discoidal cell is destitute of the insulated yellow spot, which every specimen of Po. Napi that has passed under my examination possesses.

Spring Brood


Green-veined White male first brood - Solihull 26.05.2013
Male
Photo © nfreem
Green Veined White Male (First Brood) - Chaldon, Surrey 13-April-10
Male Underside
Photo © Vince Massimo
Green-veined White female - Norfolk Estate, Sussex 14-April-2011
Female
Photo © Sussex Kipper
Green Veined White Female (First Brood) - Crawley, Sussex 27-April-06
Female Underside
Photo © Vince Massimo

Summer Brood


Green-veined-White Lincoln 4 Sept 2010- 03C6490
Male
Photo © IainLeach
Green-veined White (male), Steyning (22 August 2011)
Male Underside
Photo © Mark Colvin
Green-veined white female summer brood - Oversley Wood 21.07.2012
Female
Photo © nfreem
Green-veined White (female), Steyning (22 August 2011)
Female Underside
Photo © Mark Colvin

Subspecies: Pieris napi thomsoni

This subspecies was first defined in Warren (1968) as shown here (type locality: Sheriffmuir, West Perthshire). This subspecies exhibits minor colour differences with the subspecies sabellicae. This subspecies is found in Scotland, with most records coming from a belt between Argyllshire in the west and Fifeshire and East Lothian in the south-east. Another stronghold is the Thurso district of Caithness. In southern Scotland and in the north of England, there is no clear dividing line between the distribution of this subspecies and that of the subspecies sabellicae. This subspecies also exhibits differences between spring and summer broods, as described for the subspecies sabellicae. This subspecies differs from the subspecies sabellicae as follows:

1. On the upperside, the veins of the forewings, and sometimes hindwings, are heavily suffused with black. This is especially true in the female.

2. On the underside of the hindwings, a small proportion of adults have a yellow, rather than white, ground colour, which may be tinged with orange. This is especially true in the female.

3. The underside is generally darker, with the dark scaling on the veins appearing more black than grey.

Pieris napi thomsoni

Turning to the superficial features of the Scottish race, the most conspicuous are ... the markings of the females [where] ... the nervures of the forewings, and sometimes hindwings also, on the upperside are outlined in black, lightly or heavily. In the Scottish race many have the areas between the nervures also suffused with black. They most often have a white ground-colour and a few specimens can resemble P. napi exactly. Some however (estimated at about 25% by Mr. Thomson), have a yellow ground-colour. In these the dark suffusion can occur as in the white specimens.

A further character ... is the colouring of the under side. This mostly is darker than in normal P. napi, the dark scaling on the nervures is more black than grey. There is also a not infrequent tendency for the marking on the hindwing to spread on each side of the nervures, especially on the basal half of the wing. The ground-colour is often a deeper yellow, especially in the females, with a tinge of orange.

The male holotype and female allotype and one male and female paratype (the latter being a yellow one), are from Sheriffmuir, Dunblane, Perthshire, at 600 feet altitude; and one male and one female paratype from Dunblane at 200 feet altitude; and 11 paratypes ... of which two males and four females come from the Carron Valley, Stirlingshire, two males and two females from the Kilsyth Hills, Stirlingshire, and one female from St. Andrews, Fifeshire.

Spring Brood


Male
Green-veined White - East Lothian, Scotland 30-March-2012
Male Underside
Photo © NickMorgan
Green-Veined White - imago - Loch Cre - Unknown date [Adrian Riley]
Female
Photo © Adrian Riley
Female Underside

Summer Brood


Male
Green-veined White - East Lothian, Scotland 8-Aug-2012
Male Underside
Photo © NickMorgan
Female
Green-veined White - East Lothian, Scotland 17-Aug-2011
Female Underside
Photo © NickMorgan

Subspecies: Pieris napi britannica

This subspecies was first defined in Müller & Kautz (1939) (type locality: Ireland). This subspecies is found throughout Ireland, where it is common. This subspecies exhibits differences between spring and summer broods, as described for the subspecies sabellicae.

Spring Brood


Green-Veined White - Male Upperside 26/04/2007, Mountstewart, Co Down, Northern Ireland
Male
Photo © Dave McCormick
Male Underside
Green-Veined White - imago - Lough Georg - Unknown date [Adrian Riley]
Female
Photo © Adrian Riley
Female Underside

Summer Brood


Male
Male Underside
Female
Female Underside

Phenology  

First-brood adults typically emerge in late April, peaking around the middle of May and gradually tailing off through June. The second brood, which is always stronger than the first brood, starts to emerge in early July. However, in good years, the second brood may emerge in late June and give rise to a third brood.

Subspecies: Pieris napi sabellicae


Subspecies: Pieris napi thomsoni


Subspecies: Pieris napi britannica


The chart(s) above have been correlated with the phenology plot below, taken from the UK Butterfly Monitoring Scheme. The blue line gives average counts over the full data set from 1976 to date, and the red line gives the average for the last year.


Habitat  

The butterfly can be found in a variety of locations, including parks, gardens, meadows, woodland rides, hedgerows and, in fact, anywhere foodplants and nectar sources exist. This species favours damp areas but can also be found in small sheltered pockets, such as patches of scrub, in dry and open habitat such as chalk grassland.

Larval Foodplants  

The primary larval foodplants are Charlock (Sinapis arvensis), Cuckooflower (Cardamine pratensis), Garlic Mustard (Alliaria petiolata), Hedge Mustard (Sisymbrium officinale), Large Bitter-cress (Cardamine amara), Water-cress (Rorippa nasturtium-aquaticum), Wild Cabbage (Brassica oleracea) and Wild Radish (Raphanus raphanistrum). Crucifers (various) (Cruciferae family (various)) and Nasturtium (Tropaeolum majus) are also used.

Nectar Sources  

Adults feed primarily on Betony (Stachys officinalis), Bluebell (Hyacinthoides non-scriptus), Bugle (Ajuga reptans), Buttercups (Ranunculus spp.), Cuckooflower (Cardamine pratensis), Fleabane (Pulicaria dysenterica), Greater Stitchwort (Stellaria holostea), Ragged Robin (Lychnis flos-cuculi), Ragwort (Senecio jacobaea), Red Campion (Silene dioica), Thistles (Cirsium spp. and Carduus spp.) and Vetches (Vicia spp.).

Imago  

As in most species, the males emerge a few days earlier than the females, and spend much of their time searching for a mate as they fly along woodland rides and hedgerows. As well as taking nectar, males may also be seen congregating on mud or other surfaces that are rich in nutrients, giving rise to the phenomenon known as “mud puddling”.

As in many whites, an already-mated female will indicate an unwillingness to mate by holding her wings flat and her abdomen upright, making it impossible for a male to mate with her. However, this doesn’t always work, and females have been known to mate more than once, although this is completely unnecessary for the fertilisation of the eggs.

Subspecies: Pieris napi sabellicae

Spring Brood


Green Veined White Female (First Brood) - Crawley, Sussex 27-April-06
Photo © Vince Massimo
27-Apr-2006
Green-veined White female (1st brood) - Crawley, Sussex 17-May-06
Photo © Vince Massimo
17-May-2006
Green Veined White Male (First Brood) - Crawley, Sussex 13-April-09
Photo © Vince Massimo
13-Apr-2009
Green-Veined White - imago - Hartslock - 10-May-09 (1)
Photo © Pete Eeles
10-May-2009
Green-Veined White - imago - Midgham Lakes - 14-Apr-09 (1)
Photo © Pete Eeles
14-Apr-2009
Green-Veined White - imago - Clayton, Staffordshire - 21-Apr-09 (2) [John Wood]
Photo © John Wood
Green-Veined White - imago - Clayton, Staffordshire - 21-Apr-09 [John Wood]
Photo © John Wood
Green-veined Whites Clive Burrows [Clive Burrows]
Photo © Clive Burrows
Green-veined White - imago - Botany Bay - 14-May-10 (1)
Photo © Pete Eeles
14-May-2010
Green Veined White Male (First Brood) - Chaldon, Surrey 13-April-10
Photo © Vince Massimo
13-Apr-2010
Green-veined-White Lincoln 10 April 2011- 03C9279
Photo © IainLeach
Green-veined-White Lincoln 10 April 2011- 03C9181
Photo © IainLeach
Green-veined White female, 1st brood - Mill Road Cemetery, Cambridge 20 April 2012
Photo © NickB
Green-Veined White - imago - Thatcham - 18-Apr-12 (2) [REARED]
Photo © Pete Eeles
Green-veined White - imago - Thatcham - 30-Apr-12 (1) [REARED]
Photo © Pete Eeles
Green-veined White - imago - Woolhampton Gravel Pits - 21-Apr-12
Photo © Pete Eeles
Green Veined White - Oxfordshire May 20th 2012.
Photo © Nigel Kiteley
20-May-2012
Green-veined White female - Norfolk Estate, Sussex 14-April-2011
Photo © Sussex Kipper
15-Apr-2011
Green-veined White male, Springhead Hill, Sussex 7-May-2012
Photo © Sussex Kipper
07-May-2012
Green-veined White male (first brood) - Addington, Surrey 21-May-2012
Photo © Vince Massimo
21-May-2012
Green-veined White female (1st brood) - Crawley, Sussex 17-May-06
Photo © Vince Massimo
17-May-2006
Green-veined White male first brood - Solihull 04.05.2013
Photo © nfreem
04-May-2013
Green-veined white, 16 May 2013, Wrecclesham
Photo © Pauline
16-May-2013
Green-veined White female - Ashford, Kent 11-May-2013
Photo © Hoggers
11-May-2013
Green-veined White male first brood - Solihull 26.05.2013
Photo © nfreem
26-May-2013
Green-veined White Mating Pair with Female Onlooker Debdon Forest, Northumberland 26-05-2013
Photo © Graham Beckwith
26-May-2013

Summer Brood


Green-Veined White - imago - Crabtree Plantation - 03-Jul-06 (0443)
Photo © Pete Eeles
03-Jul-2006
Green-Veined White - imago -  Macclesfield Forest - 11-Aug-07 [Janette Bowler]
Photo © Janette Bowler
Green-veined White - imago - Fakenham - Unknown date [Adrian Riley]
Photo © Adrian Riley
Green Veined White Female (Summer Brood - Reared) - Caterham, Surrey 6-July-10
Photo © Vince Massimo
06-Jul-2010
Green-veined White
Photo © Charlotte Brett
Green-veined White - imago - Stockbridge Down - 20-Jul-10 (1)
Photo © Pete Eeles
20-Jul-2010
Green Veined White Male (Second Brood) - Southwater Wood, Sussex 5-July-10
Photo © Vince Massimo
05-Jul-2010
Green Veined White Male (Second Brood) - Woldingham, Surrey 23-June-11
Photo © Vince Massimo
23-Jun-2011
Green Veined White Pair (Second Brood) - Woldingham, Surrey 30-June-11
Photo © Vince Massimo
30-Jun-2011
Green-veined White female (third brood) - Stanwell Moor (Middx) 19 Nov 2011
Photo © millerd
19-Nov-2011
Green-veined White female (third brood) - Stanwell Moor (Middx) 19 Nov 2011
Photo © millerd
19-Nov-2011
Green-veined White Female (Third Brood) - Chaldon, Surrey 15-Sept-11
Photo © Vince Massimo
15-Sep-2011
Green-Veined White - imago - Pamber Forest - 24-Jun-11 (1)
Photo © Pete Eeles
Green-veined White (female), Steyning (22 August 2011)
Photo © Mark Colvin
22-Aug-2011
Green-veined White (male), Steyning (22 August 2011)
Photo © Mark Colvin
22-Aug-2011
Green-veined-White Lincoln 14 August 2011- 03C9501
Photo © IainLeach
Green-veined-White- Lincoln 14 August 2011 03C8282
Photo © IainLeach
Green-veined-White Lincoln 12 August 2010- 03C7401
Photo © IainLeach
Green-veined-White Lincoln 4 Sept 2010- 03C6490
Photo © IainLeach
Green-veined-White Lincoln 6 August 2011- 03C6088
Photo © IainLeach
Green-veined white female summer brood - Oversley Wood 21.07.2012
Photo © nfreem
21-Jul-2012
Green-veined White male (Second Brood) - Chaldon, Surrey 30-July-2012
Photo © Vince Massimo
30-Jul-2012

Subspecies: Pieris napi thomsoni

Spring Brood


Green-veined White - East Lothian, Scotland 30-March-2012
Photo © NickMorgan
Green-Veined White - imago - Loch Cre - Unknown date [Adrian Riley]
Photo © Adrian Riley
Green-Veined White - imago - North Kessock - Unknown date [Phil Brown]
Photo © Phil Brown
Green-veined White female - Binning Wood, East Lothian 16/05/2012
Presuming that this is ssp. thomsoni according to the distribution map.
Photo © NickMorgan

Summer Brood


Green-veined White - East Lothian, Scotland 8-Aug-2012
Photo © NickMorgan
Green-veined White - East Lothian, Scotland 17-Aug-2011
Photo © NickMorgan
17-Aug-2011

Subspecies: Pieris napi britannica

Spring Brood


Green-Veined White - Male Upperside 26/04/2007, Mountstewart, Co Down, Northern Ireland
Photo © Dave McCormick
26-Apr-2007
Green-Veined White - Female - 09/05/2007, Mountstewart, Co Down, Northern Ireland
Photo © Dave McCormick
09-May-2007
Green-Veined White - Mating Pair 11/05/2008 - Whitespots Country Park, Co Down, Northern Ireland
Photo © Dave McCormick
11-May-2008
Green-Veined White - imago - Lough Georg - Unknown date [Adrian Riley]
Photo © Adrian Riley

Summer Brood

Aberrations  

This species exhibits great variation in the colour and shade of the ground colour as well as the extent of the spotting. As usual in the 'whites' many aberrations are unique to either male or female specimens, and being seasonally dimorphic some aberrations are also specific to the generation or 'brood'.

This species has been the subject of much experimental breeding in the past, particularly of the impressive aberrations suphurea and fasciata. The aberration sulphurea has been recorded chiefly in Scotland and Ireland and specimens from these areas were bred en masse by entomologists in the early 20th century. These aberrations were popular with breeders and collectors and while scarce now in the wild, a large number of these impressive forms can be found in collections.

There are 79 named aberrations known to occur in Britain. Click here to see a full list of aberrations for this species.

Ovum  

Eggs are laid singly on the underside of a leaf of the foodplant. Small plants are preferred, and the eggs hatch in approximately a week depending on temperature. Eggs are sometimes found on the same plants as those used by the Orange-tip. However, the two species are not in competition since the Green-veined White eats the leaves of the plant, whereas the Orange-tip eats the developing seed pods.


Green-veined White - Bristol - 21-May-09
Photo © Denise
Green-Veined White - ovum - Thatcham - 12-May-09 (1)
Photo © Pete Eeles
12-May-2009
Green-Veined White - ovum - Unknown location - 2003 [REARED] [Reg Fry]
Photo © Reg Fry
Green-Veined White - ovum - Woolhampton Gravel Pits - 14-Sep-11 (1)
Photo © Pete Eeles

Larva  

The larva eats its eggshell on hatching, before starting to feed on the leaves of the foodplant. There are 4 moults in total and this stage lasts between 3 and 4 weeks.


Green-veined White, (early instar larva) - Bristol - 28-May-09
Photo © Denise
Green-Veined White - larva - Thatcham - 16-May-09 (1)
Photo © Pete Eeles
16-May-2009
Green-Veined White - larva - Thatcham - 06-Jun-09 (2) {REARED}
Photo © Pete Eeles
06-Jun-2009
Green-Veined White - larva - Thatcham - 25-May-09 (1)
Photo © Pete Eeles
25-May-2009
Green Veined White Larva - Caterham, Surrey 27-June-10
Photo © Vince Massimo
27-Jun-2010
Green-Veined White - larva - Thatcham - 04-Oct-11 (1) {REARED}
Photo © Pete Eeles
Green-Veined White - larva - Thatcham - 04-Oct-11 (2) {REARED}
Photo © Pete Eeles

Pupa  

The pupa is generally formed away from the foodplant, low down in vegetation. It is supported by a silk girdle and the cremaster. The pupa has 2 colour forms, green and light brown, although some pupae have an intermediate colouring. This stage lasts around 10 days unless overwintering.


Green-Veined White - pupa - Unknown location - 2003 [REARED] [Reg Fry]
Photo © Reg Fry
Green Veined White Pupa (Reared - 2 Days Before Hatching) - Caterham, Surrey 4-July-10
Photo © Vince Massimo
04-Jul-2010
Green Veined White Pupa Female (Reared - 2 Hours Before Hatching) - Caterham, Surrey 6-July-10
Photo © Vince Massimo
06-Jul-2010
Green-Veined White - pupa - Thatcham - 07-Oct-11 (1) {REARED}
Photo © Pete Eeles
Green-veined White pupa (ssp. thomsoni) - East Lothian, Scotland 5-Jan-2012
ssp. thomsoni
Photo © NickMorgan
Green-veined White pupa (ssp. thomsoni) - East Lothian, Scotland 5-Jan-2012
ssp. thomsoni
Photo © NickMorgan

Similar Species  

Large White

Description to be completed.

Orange-tip

Description to be completed.

Small White

The Green-veined White and Small White are most easily distinguished by their undersides, where the Green-veined White has pronounced markings along the veins which are absent in the Small White.


Green-veined White (left) and Small White (right)

It is much more difficult to distinguish between the Green-veined White and Small White based on the upperside, since the amount of marking is highly variable. In general, the veins of the Green-veined White are more pronounced. Also, the marking at the apex of the forewing of a Green-veined White often extends down the along the edge of the forewing and is not contiguous. The marking at the apex of a Small White never extends down the edge of the forewing and is unbroken.


Green-veined White male (left) and Small White male (right)
Videos  

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Sites  

Click here to see the distribution of this species overlaid with specific site information. Alternatively, select one of the sites listed below.

Sites
Aberffraw Dunes, Arthur's Seat, Ashampstead Common, Banstead Woods, Bedfont Lakes Country Park LNR, Bryncelyn Hall, Cuerden Valley Park, Devil's Ditch, Durlston NNR, Eas na Circe, Fleam Dyke, Glasdrum Wood, Glen Loy, Glenarm, Higher Hyde, Horsenden Hill, Hounslow Heath LNR, Kinghorn Loch Path, Latton Woods, Lavernock, Linn Dean, Mansmead wood, Meanwood Park, Midgham Lakes, Mill Hill, Moors Valley Country Park, Morgaston Wood, Moss Field, Mynydd Marian, Nupend Wood, Old Down, Basingstoke, Rookery, Roudsea Wood NNR, Ryton Woods Meadows, Spean Bridge, Strumpshaw Fen, Sutton Bingham Reservoir, Thurso, Tophill Low, Viking Field/LesleySears, Winsdon Hill

Conservation Status  

This is the most widespread butterfly in the British Isles and its status is considered relative-stable. It is not, therefore, a species of conservation concern.

UK BAP StatusDistribution TrendPopulation Trend
Not ListedStableStable

From The State of Butterflies in Britain and Ireland and the UK Biodiversity Action Plan (UK BAP) (2007 review).


Links  

The following links provide additional information on this butterfly.

References  

The species description provided here has been derived from the author's own observations and the information contained in the following works:

  • Asher (2001). Asher, J., Warren, M., Fox, R., Harding, P., Jeffcoate, G. and Jeffcoate, S.: The Millennium Atlas of Butterflies in Britain and Ireland. 2001.
  • Fox (2006). Fox, R., Asher, J., Brereton, T., Roy, D. and Warren, M.: The State of Butterflies in Britain and Ireland. 2006.
  • Frohawk (1914). Frohawk, F.W.: A Natural History of British Butterflies. 1914.
  • Howarth (1973). Howarth, T.G.: South's British Butterflies. 1973.
  • Nash (2012). Nash, D., Boyd, T. and Hardiman, D.: Ireland's Butterflies: A Review. 2012.
  • Riley (2007). Riley, A.M.: British and Irish Butterflies: The Complete Identification, Field and Site Guide to the Species, Subspecies and Forms. 2007.
  • South (1906). South, R.: The Butterflies Of The British Isles. 1906.
  • Thomas & Lewington (2010). Thomas, J. and Lewington, R.: The Butterflies of Britain and Ireland. Edn.2. 2010.

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