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Butterfly taxonomy (classification) The skippers The swallowtails The whites The hairstreaks, coppers and blues. Includes the Duke of Burgundy. The nymphalids, fritillaries and browns. Includes the Monarch.
Clouded Yellow. 25/09/2009. Beachy Head, Sussex.
Wingspan
Male: 52 - 58mm
Female: 54 - 62mm
Photo © badgerbob
Clouded Yellow

Colias croceus
Pronunciation KO-lee-uss
KROH-see-uss
B&F No.: 1545
Superfamily: Papilionoidea
Family: Pieridae
Subfamily: Coliadinae
Genus: Colias
Subgenus:  
Species: croceus (Geoffroy, 1785)
Subspecies:croceus (Geoffroy, 1785)
Form:helice (Hübner, 1779)
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Introduction  

The Clouded Yellow is primarily an immigrant to the UK, originating from north Africa and southern Europe, with numbers varying greatly from year to year – an estimated 36,000 butterflies appearing in one of the infrequent “Clouded Yellow” years in 1947. In more recent years, it has been shown that this species has successfully overwintered in the south of England. However, it is believed that the majority of individuals perish, since both larva and pupa of this continuously-brooded species are easily killed by damp and frost. In good years this species can produce up to 3 generations in the UK. In flight, this species is often mistaken for one of the commoner “whites”, but the orange-yellow colour is quite distinctive, even in flight, and unlike any other species. The Clouded Yellow has a distribution befitting a highly-migratory species, and can be found anywhere in the British Isles. Many immigrants remain near the coast where they feed, mate, and lay eggs. Others disperse inland and this species is found in both Scotland and Ireland in good years.

Subspecies: Colias croceus croceus

The nominate subspecies was first defined in Geoffroy (1785) as shown here (type locality: Paris, France). The British population is represented by this subspecies.


Clouded Yellow, male - Mill Hill, West Sussex - 12.9.2010 [Colin Knight] 7
Male
Photo © Colin Knight
Clouded Yellow Male - Birling Gap, Sussex 1-Oct-09
Male Underside
Photo © Vince Massimo
Clouded Yellow Female - Oreshari Meadows, nr Arda Valley, Bulgaria June 2012
Female
Photo © Paul Wetton
Clouded Yellow, Amberley, 6 September, 2007
Female Underside
Photo © Sussex Kipper

Form: Colias croceus croceus f. helice

This form was first defined in Hübner (1779) as shown here and as shown in this plate (type locality: Not stated). The form helice occurs in the female, where the individual is a creamy white, rather than yellow, in colour.


Male
Male Underside
Female
Clouded Yellow - imago - Swanwick - 30-Jul-05 [Andy Collins]
Female Underside
Photo © Andy Collins

Phenology  

The first immigrants of this species start to arrive to our shores in late May or early June, with much larger numbers appearing in July and August, as the offspring of the first arrivals mingle with new immigrants.

Subspecies: Colias croceus croceus


The chart(s) above have been correlated with the phenology plot below, taken from the UK Butterfly Monitoring Scheme. The blue line gives average counts over the full data set from 1976 to date, and the red line gives the average for the last year.


Habitat  

This butterfly can be found in just about any open habitat in the countryside, including coastal cliffs, open downland, and fields containing the larval foodplants of Clovers, Lucerne and Bird's-foot Trefoil.

Larval Foodplants  

The primary larval foodplants are Clovers (various) (Trifolium spp.) and Lucerne (Medicago sativa). Bird's-foot Trefoil (Lotus corniculatus) is also used.

Nectar Sources  

Adults feed primarily on Dandelion (Taraxacum agg.), Fleabane (Pulicaria dysenterica), Knapweeds (Centaurea spp.), Marjoram (Origanum vulgare), Ragwort (Senecio jacobaea), Thistles (Cirsium spp. and Carduus spp.) and Vetches (Vicia spp.).

Imago  

This strong-flying butterfly always settles with its wings closed and so the dark borders on the uppersides of the wings are only visible when in flight. There is no elaborate courtship and, having mated, the female is subsequently able to lay an extraordinary number of eggs – up to 600 have been recorded from a single female.

Subspecies: Colias croceus croceus


Clouded Yellow Female, Birling Gap, Sussex 4-Sept-07
Photo © Vince Massimo
04-Sep-2007
Clouded Yellow - Beachy Head, 27th August 2009
Photo © NickB
27-Sep-2009
Clouded Yellow pair, Beachy Head 27th Sept 2009
Photo © NickB
27-Sep-2009
Clouded Yellow Pair - Birling Gap, Sussex 1-Oct-09
Photo © Vince Massimo
01-Oct-2009
Clouded Yellow Female - Birling Gap, Sussex 1-Oct-09
Photo © Vince Massimo
01-Oct-2009
Clouded Yellow Male - Birling Gap, Sussex 1-Oct-09
Photo © Vince Massimo
01-Oct-2009
Clouded Yellow, Amberley, 6 September, 2007
Photo © Sussex Kipper
06-Sep-2007
Clouded Yellow, male - Mill Hill, West Sussex - 12.9.2010 [Colin Knight] 3
Photo © Colin Knight
12-Sep-2010
Clouded Yellow, male - Mill Hill, West Sussex - 12.9.2010 [Colin Knight] 5
Photo © Colin Knight
Clouded Yellow, male - Mill Hill, West Sussex - 12.9.2010 [Colin Knight] 7
Photo © Colin Knight
12-Sep-2010
Clouded Yellow pair - Sora, Slovenia September 2012
Photo © Simon2
22-Sep-2012
Clouded Yellow (female), Kithurst Hill, Sussex (7 September 2011)
Photo © Mark Colvin
07-Sep-2011
Clouded Yellow (m) Birling Gap, Sussex  25th September 2009
Photo © millerd
25-Sep-2009
Clouded Yellow (m) Birling Gap, Sussex  25th September 2009
Photo © millerd
25-Sep-2009
Clouded Yellow (mating pair) Birling Gap, Sussex  25th September 2009
Photo © millerd
25-Sep-2009
Clouded Yellow (m) close-up Birling Gap, Sussex  25th September 2009
Photo © millerd
25-Sep-2009
Clouded Yellow. Seaford. 11/8/2009.
Photo © badgerbob
11-Aug-2009
Clouded Yellow (Colias croceus), female, - Velingrad, SW Bulgaria (07/06/12)
Photo © Michaeljf
Clouded Yellow Female - Oreshari Meadows, nr Arda Valley, Bulgaria June 2012
Photo © Paul Wetton
Clouded Yellow. 25/09/2009. Beachy Head, Sussex.
Photo © badgerbob
25-Sep-2009
Clouded Yellow - Cissbury, Sussex 18-Oct-2011
Photo © Sussex Kipper
18-Oct-2011
Clouded Yellow female - Birling Gap, Sussex 4-Sept-07
Photo © Vince Massimo
04-Sep-2007
Clouded Yellow male - Birling Gap, Sussex 1-Oct-09
Photo © Vince Massimo
01-Oct-2009

Form: Colias croceus croceus f. helice


Clouded Yellow - form - Thatcham - 12-Nov-07 (1222) {REARED}
Photo © Pete Eeles
12-Nov-2007
Clouded Yellow Female (f.helice) - Birling Gap, Sussex 1-Oct-09
Photo © Vince Massimo
01-Oct-2009
Clouded Yellow Female (f.helice) - Birling Gap, Sussex 28-Sept-09
Photo © Vince Massimo
28-Sep-2009
Clouded Yellow - imago - Swanwick - 30-Jul-05 [Andy Collins]
Photo © Andy Collins
Clouded Yellow - imago - Sarnano, Italy - 17-Jun-08 (1)
Photo © Pete Eeles
17-Jun-2008
Clouded Yellow f.helice  Birling Gap, Sussex  25th September 2009
Photo © millerd
25-Sep-2009
Clouded Yellow - form - Carymoor - 03-Aug-06 [Jules Cross]
Photo © Jules Cross
03-Aug-2006
Clouded Yellow - imago - Lake Kerkini, Greece - 08-Jun-09 (1)
Photo © Pete Eeles
08-Jun-2009
Clouded Yellow f.helice  Birling Gap, Sussex  25th September 2009
Photo © millerd
25-Sep-2009
Clouded Yellow ab.helice. 13/10/09. Seaford, Sussex.
Photo © badgerbob
13-Oct-2009

Aberrations  

Description to be completed.

Ovum  

The skittle-shaped eggs are laid singly on the upperside of leaves of the foodplant. They are pale yellow when first laid, turning orange prior to hatching, and hatch in about a week.


Clouded Yellow ovum, Menorca
Photo © NickMorgan
Clouded Yellow - ovum - Portugal - 13-Sep-06 [Colin Baker]
Photo © Colin Baker
Clouded Yellow eggs - Rhone Valley, Switzerland 20-Oct-2012
Photographed within minutes of being laid, in Switzerland on 20th October 2012
Photo © padfield
20-Oct-2012
Clouded Yellow - ovum - Thatcham - 17-Jul-12 (1) [REARED]
Photo © Pete Eeles

Larva  

The rate of growth of the larva is greatly dependent on conditions, especially temperature. The duration of this stage is therefore highly variable, ranging from 3 to 6 weeks. There are 4 moults in total.


Clouded Yellow 1st instar larva. Bournemouth 12.Oct 06
Photo © Mikhail
Clouded Yellow - larva - Thatcham - 02-Nov-07 (2) {REARED}
Photo © Pete Eeles
02-Nov-2007
Clouded Yellow - larva - Thatcham - 26-Oct-07 (1) {REARED}
Photo © Pete Eeles
26-Oct-2007
Clouded Yellow - larva - Thatcham - 31-Oct-07 (1224) {REARED}
Photo © Pete Eeles
31-Oct-2007
Clouded Yellow - larva - Thatcham - 31-Oct-07 (1225) {REARED}
Photo © Pete Eeles
31-Oct-2007
Clouded Yellow - larva - Thatcham - 31-Oct-07 (1226) {REARED}
Photo © Pete Eeles
31-Oct-2007

Pupa  

The pupa is attached to a foodplant stem by a silk girdle and the cremaster. This stage lasts 2 or 3 weeks.


Clouded Yellow pupa. Bournemouth 6 Jul 01
Photo © Mikhail
Clouded Yellow - pupa - Thatcham - 06-Nov-07 (1) {REARED}
Photo © Pete Eeles
06-Nov-2007
Clouded Yellow - pupa - Thatcham - 25-Nov-07 (2) {REARED}
Photo © Pete Eeles
25-Nov-2007
Clouded Yellow - pupa - Unknown location - Unknown date [REARED] [Adrian Dexter]
Photo © Adrian Dexter
Clouded Yellow - pupa - Thatcham - 31-Aug-12 (1) [REARED]
Photo © Pete Eeles

Similar Species  

Berger's Clouded Yellow

Of the three species of Clouded Yellow found in the British Isles, the Clouded Yellow is both the commonest and the easiest to identify. When in flight, the orange appearance of the Clouded Yellow is unlike any other British butterfly. When settled, the lemon-coloured underside of the Clouded Yellow allows us to distinguish this species from Berger's Clouded Yellow which has a much paler underside. This diagnostic holds true even in the helice form of female Clouded Yellow where the orange colouring is replaced by a creamy white.


1. Clouded Yellow 2. Clouded Yellow (f. helice)
3. Berger's Clouded Yellow (male) 4. Berger's Clouded Yellow (female)

Pale Clouded Yellow

Of the three species of Clouded Yellow found in the British Isles, the Clouded Yellow is both the commonest and the easiest to identify. When in flight, the orange appearance of the Clouded Yellow is unlike any other British butterfly. When settled, the lemon-coloured underside of the Clouded Yellow allows us to distinguish this species from Pale Clouded Yellow which has a much paler underside. This diagnostic holds true even in the helice form of female Clouded Yellow where the orange colouring is replaced by a creamy white.


1. Clouded Yellow 2. Clouded Yellow (f. helice)
3. Pale Clouded Yellow (male) 4. Pale Clouded Yellow (female)
Videos  

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Sites  

Click here to see the distribution of this species overlaid with specific site information. Alternatively, select one of the sites listed below.

Sites
Banstead Woods, Durlston NNR, Mill Hill, Moss Field, Old Winchester Hill, West Yatton Down

Conservation Status  

The Clouded Yellow is one of the most-widespread species in Europe and is not a species of conservation concern.

UK BAP StatusDistribution TrendPopulation Trend
Not ListedStableLarge Increase

From The State of Butterflies in Britain and Ireland and the UK Biodiversity Action Plan (UK BAP) (2007 review).


Links  

The following links provide additional information on this butterfly.

References  

The species description provided here has been derived from the author's own observations and the information contained in the following works:

  • Asher (2001). Asher, J., Warren, M., Fox, R., Harding, P., Jeffcoate, G. and Jeffcoate, S.: The Millennium Atlas of Butterflies in Britain and Ireland. 2001.
  • Fox (2006). Fox, R., Asher, J., Brereton, T., Roy, D. and Warren, M.: The State of Butterflies in Britain and Ireland. 2006.
  • Frohawk (1914). Frohawk, F.W.: A Natural History of British Butterflies. 1914.
  • Howarth (1973). Howarth, T.G.: South's British Butterflies. 1973.
  • Nash (2012). Nash, D., Boyd, T. and Hardiman, D.: Ireland's Butterflies: A Review. 2012.
  • Riley (2007). Riley, A.M.: British and Irish Butterflies: The Complete Identification, Field and Site Guide to the Species, Subspecies and Forms. 2007.
  • South (1906). South, R.: The Butterflies Of The British Isles. 1906.
  • Thomas & Lewington (2010). Thomas, J. and Lewington, R.: The Butterflies of Britain and Ireland. Edn.2. 2010.

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