As explained to me by GP during several discussions over the years, lately included.. thought I would collage 3 pics all taken in the Alps this year showing trends between Alpine and Darwin's Heath. Poor quality given the 800 pixel limit but open for comments if any!
gardarwiniana
gardarwiniana
My own webpages.... http://ptkbutterflies.wix.com/photo-art
- Padfield
- Administrator
- Posts: 8373
- Joined: Sun Nov 05, 2006 10:19 pm
- Location: Leysin, Switzerland
- Contact:
Re: gardarwiniana
Nice!
You could perhaps add a picture of arcania as a fourth on the right ...

It continues the sequence.
As you know, my theory is that gardetta was the mountain form of arcania during the last Interglacial. They were separated and went their own ways during the subsequent Ice Age, then met again in the present Interglacial, when hybridisation between what were now different species produced darwiniana. There are regions in France where all three taxa are known to hybridise ...
I have absolutely no evidence to back up the theory, but there is clearly a close relationship between these three taxa, with darwiniana in the middle. Leander fits into the picture somehow, too!
Guy
You could perhaps add a picture of arcania as a fourth on the right ...

It continues the sequence.
As you know, my theory is that gardetta was the mountain form of arcania during the last Interglacial. They were separated and went their own ways during the subsequent Ice Age, then met again in the present Interglacial, when hybridisation between what were now different species produced darwiniana. There are regions in France where all three taxa are known to hybridise ...
I have absolutely no evidence to back up the theory, but there is clearly a close relationship between these three taxa, with darwiniana in the middle. Leander fits into the picture somehow, too!
Guy
Guy's Butterflies: https://www.guypadfield.com
The Butterflies of Villars-Gryon : https://www.guypadfield.com/villarsgryonbook.html
The Butterflies of Villars-Gryon : https://www.guypadfield.com/villarsgryonbook.html
- Pete Eeles
- Administrator & Stock Contributor
- Posts: 6869
- Joined: Tue Jan 17, 2006 6:10 pm
- Location: Thatcham, Berkshire
- Contact:
Re: gardarwiniana
Nice visualisation Paul and very reminiscent of Large Heath (scotica -> polydama -> davus). I guess the jury's out on the taxonomic status in this case!
Cheers,
- Pete

Cheers,
- Pete
Life Cycles of British & Irish Butterflies: http://www.butterflylifecycles.com
British & Irish Butterflies Rarities: http://www.butterflyrarities.com
British & Irish Butterflies Rarities: http://www.butterflyrarities.com
- Padfield
- Administrator
- Posts: 8373
- Joined: Sun Nov 05, 2006 10:19 pm
- Location: Leysin, Switzerland
- Contact:
Re: gardarwiniana
Most authors seem to regard gardetta and darwiniana as conspecific these days. This is what the Swiss Bible has to say about their distribution (it treats them as separate species, but was written in 1987):
'The species [darwiniana] has pure populations in the southern valleys of the Alps, notably in the Tessin, in the Val Mesolcina and on the southern slope of the Simplon. Visibly mixed populations are found practically everywhere in the southern valleys where C. gardetta and C. darwiniana meet, as well as in the Haut Valais, in the valleys of the Alpes uranaises and to the north of the passes of San Bernadino, La Bernina and Il Fuorn. These transitional forms, designated f. philedarwiniana and visibly hybrids, are often confused with C. darwiniana and sometimes with C. arcania. Their distribution needs to be studied in greater depth'.
Darwin himself would love all this evidence of evolution in action!
Guy
'The species [darwiniana] has pure populations in the southern valleys of the Alps, notably in the Tessin, in the Val Mesolcina and on the southern slope of the Simplon. Visibly mixed populations are found practically everywhere in the southern valleys where C. gardetta and C. darwiniana meet, as well as in the Haut Valais, in the valleys of the Alpes uranaises and to the north of the passes of San Bernadino, La Bernina and Il Fuorn. These transitional forms, designated f. philedarwiniana and visibly hybrids, are often confused with C. darwiniana and sometimes with C. arcania. Their distribution needs to be studied in greater depth'.
Darwin himself would love all this evidence of evolution in action!
Guy
Guy's Butterflies: https://www.guypadfield.com
The Butterflies of Villars-Gryon : https://www.guypadfield.com/villarsgryonbook.html
The Butterflies of Villars-Gryon : https://www.guypadfield.com/villarsgryonbook.html
- Pete Eeles
- Administrator & Stock Contributor
- Posts: 6869
- Joined: Tue Jan 17, 2006 6:10 pm
- Location: Thatcham, Berkshire
- Contact:
Re: gardarwiniana
Indeed - and so do Ipadfield wrote:Darwin himself would love all this evidence of evolution in action!

Cheers,
- Pete
Life Cycles of British & Irish Butterflies: http://www.butterflylifecycles.com
British & Irish Butterflies Rarities: http://www.butterflyrarities.com
British & Irish Butterflies Rarities: http://www.butterflyrarities.com
Re: gardarwiniana
Hi,
A little late with the reply, but thought you might find this interesting:
Martin Wiemers (1998). Coenonympha darwiniana - a hybrid taxon? New insights through allozyme electrophoresis
http://www.univie.ac.at/population-ecol ... s_1998.pdf
All the evidence you need to back your theory, Guy...
Cheers,
A little late with the reply, but thought you might find this interesting:
Martin Wiemers (1998). Coenonympha darwiniana - a hybrid taxon? New insights through allozyme electrophoresis
http://www.univie.ac.at/population-ecol ... s_1998.pdf
All the evidence you need to back your theory, Guy...

Cheers,
Nikolay Shtinkov (http://nsbutterflies.com)
- Padfield
- Administrator
- Posts: 8373
- Joined: Sun Nov 05, 2006 10:19 pm
- Location: Leysin, Switzerland
- Contact:
Re: gardarwiniana
Fascinating! Thank you, Nik.
The scenario on p.58 is exactly what I had speculated, albeit with considerably more flesh on the bones.
Guy
The scenario on p.58 is exactly what I had speculated, albeit with considerably more flesh on the bones.
Guy
Guy's Butterflies: https://www.guypadfield.com
The Butterflies of Villars-Gryon : https://www.guypadfield.com/villarsgryonbook.html
The Butterflies of Villars-Gryon : https://www.guypadfield.com/villarsgryonbook.html